1988-1992:     PhD at Vrije Universiteit, Amsterdam, The Netherlands; Thesis: "Alternative splicing generates neuropeptide diversity in Lymnaea stagnalis"

1992-1993:     Post-doctoral fellow, Utrecht University, Utrecht, The Netherlands

1993-present:  Assistant professor, Reproductive biology group, Developmental Biology, Department of Biology, Faculty of Science, Utrecht Univerity, Utrecht, The Netherlands