| 1980-1986 | School of Chemistry | Utrecht University, NL |
| 1986-1991 | Graduate Study at the CBLE | Utrecht University, NL |
| 1991-1994 | Post-doctoral research fellow | MSKCC (New York USA) |
| 1994-1996 | Post-doctoral research fellow | Heidelberg, Germany |
| 1997-2002 | Junior group leader | Heidelberg, Germany |
| 2002- | Professor of veterinary biochemistry | Utrecht University, NL |
| 2004-2020 | Chairman, Dept. Biochem. and Cell Biol. | Utrecht University, NL |
| 2010- | Director, Institute of Biomembranes | Utrecht University, NL |